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This is supported by particular changes in the gene expression (i.e., increased Nos1 , Nos2 , and Nos3 expression along with the increased Ptgs2 expression) (Figure 16)

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Research has revealed a correlation between the lack of expression of LDHA, along with several other glycolytic genes such as carbonic anhydrase 9 (CA9) and VEGFA (vascular endothelial growth factor A), and the hypermethylation of these gene promoter regions in response to D-2HG (Figure 2)